4.5: Plant Roots
- Page ID
- 25122
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\(\newcommand{\avec}{\mathbf a}\) \(\newcommand{\bvec}{\mathbf b}\) \(\newcommand{\cvec}{\mathbf c}\) \(\newcommand{\dvec}{\mathbf d}\) \(\newcommand{\dtil}{\widetilde{\mathbf d}}\) \(\newcommand{\evec}{\mathbf e}\) \(\newcommand{\fvec}{\mathbf f}\) \(\newcommand{\nvec}{\mathbf n}\) \(\newcommand{\pvec}{\mathbf p}\) \(\newcommand{\qvec}{\mathbf q}\) \(\newcommand{\svec}{\mathbf s}\) \(\newcommand{\tvec}{\mathbf t}\) \(\newcommand{\uvec}{\mathbf u}\) \(\newcommand{\vvec}{\mathbf v}\) \(\newcommand{\wvec}{\mathbf w}\) \(\newcommand{\xvec}{\mathbf x}\) \(\newcommand{\yvec}{\mathbf y}\) \(\newcommand{\zvec}{\mathbf z}\) \(\newcommand{\rvec}{\mathbf r}\) \(\newcommand{\mvec}{\mathbf m}\) \(\newcommand{\zerovec}{\mathbf 0}\) \(\newcommand{\onevec}{\mathbf 1}\) \(\newcommand{\real}{\mathbb R}\) \(\newcommand{\twovec}[2]{\left[\begin{array}{r}#1 \\ #2 \end{array}\right]}\) \(\newcommand{\ctwovec}[2]{\left[\begin{array}{c}#1 \\ #2 \end{array}\right]}\) \(\newcommand{\threevec}[3]{\left[\begin{array}{r}#1 \\ #2 \\ #3 \end{array}\right]}\) \(\newcommand{\cthreevec}[3]{\left[\begin{array}{c}#1 \\ #2 \\ #3 \end{array}\right]}\) \(\newcommand{\fourvec}[4]{\left[\begin{array}{r}#1 \\ #2 \\ #3 \\ #4 \end{array}\right]}\) \(\newcommand{\cfourvec}[4]{\left[\begin{array}{c}#1 \\ #2 \\ #3 \\ #4 \end{array}\right]}\) \(\newcommand{\fivevec}[5]{\left[\begin{array}{r}#1 \\ #2 \\ #3 \\ #4 \\ #5 \\ \end{array}\right]}\) \(\newcommand{\cfivevec}[5]{\left[\begin{array}{c}#1 \\ #2 \\ #3 \\ #4 \\ #5 \\ \end{array}\right]}\) \(\newcommand{\mattwo}[4]{\left[\begin{array}{rr}#1 \amp #2 \\ #3 \amp #4 \\ \end{array}\right]}\) \(\newcommand{\laspan}[1]{\text{Span}\{#1\}}\) \(\newcommand{\bcal}{\cal B}\) \(\newcommand{\ccal}{\cal C}\) \(\newcommand{\scal}{\cal S}\) \(\newcommand{\wcal}{\cal W}\) \(\newcommand{\ecal}{\cal E}\) \(\newcommand{\coords}[2]{\left\{#1\right\}_{#2}}\) \(\newcommand{\gray}[1]{\color{gray}{#1}}\) \(\newcommand{\lgray}[1]{\color{lightgray}{#1}}\) \(\newcommand{\rank}{\operatorname{rank}}\) \(\newcommand{\row}{\text{Row}}\) \(\newcommand{\col}{\text{Col}}\) \(\renewcommand{\row}{\text{Row}}\) \(\newcommand{\nul}{\text{Nul}}\) \(\newcommand{\var}{\text{Var}}\) \(\newcommand{\corr}{\text{corr}}\) \(\newcommand{\len}[1]{\left|#1\right|}\) \(\newcommand{\bbar}{\overline{\bvec}}\) \(\newcommand{\bhat}{\widehat{\bvec}}\) \(\newcommand{\bperp}{\bvec^\perp}\) \(\newcommand{\xhat}{\widehat{\xvec}}\) \(\newcommand{\vhat}{\widehat{\vvec}}\) \(\newcommand{\uhat}{\widehat{\uvec}}\) \(\newcommand{\what}{\widehat{\wvec}}\) \(\newcommand{\Sighat}{\widehat{\Sigma}}\) \(\newcommand{\lt}{<}\) \(\newcommand{\gt}{>}\) \(\newcommand{\amp}{&}\) \(\definecolor{fillinmathshade}{gray}{0.9}\)Until now we discussed soil organisms in the animal kingdom, but soil life also includes plants. Healthy plant roots are essential for good crop yields. This is why plants evolved to expend so much energy on growing a sizable root system. During early to mid-season, corn plants send about 20 percent of the sugars produced during photosynthesis to the roots. These sugars are for root growth, and they provide material to excrete that nourishes mycorrhizal fungi and the wide variety of organisms in the rhizosphere. Roots are clearly influenced by the soil in which they live, and their extent and health are good indicators of soil health. Over eons, roots and their associated microorganisms (the root microbiome) have played an important role in capturing and storing nutrients such as calcium, magnesium and phosphorus from weathering of minerals in rocks and grains, and then making them available for plant growth. Plant roots will not grow well if the soil is compact, is low in nutrients or water, includes high populations of root pathogens, has high or low pH, or has other problems. Conversely, plant roots also influence the soil in which they grow. The physical pressure of roots growing through soil helps form aggregates by bringing particles closer together. Small roots also help bind particles together. In addition, many organic compounds are given off, or exuded, by plant roots and provide nourishment for soil organisms living on or near the roots. The zone surrounding roots is one of especially great numbers and activity of organisms that live off root exudates and sloughed-off cells. This increased activity by microorganisms, plus the slight disruption caused as roots grow through the soil, enhances the use of active (“dead”) organic matter by organisms, which increases nutrient availability to the plant. A sticky layer surrounding roots, called the mucigel, provides close contact between microorganisms, soil minerals and the plant (Figure 4.4). Root hairs, those small protrusions that grow from the outermost root layer (the epidermis), enable better access to water and nutrients by providing more contact with soil, and help form aggregates. Plant roots also contribute to organic matter accumulation. They are usually well distributed in the soil and may be slower to decompose than surface residues, even if incorporated by plowing or harrowing.
For plants with extensive root systems, such as grasses, the amount of living tissue below ground may actually be greater than the amount of leaves and stems we see above ground.
The soil population must be considered from the point of view of a biological complex; it is not sufficient to separate it into different constituent groups.
—S.A. Waksman, 1923